My general research interests include the neurobiology of learning and memory, neurocognitive aging, neuropathology, clinical neuropsychology, and animal behavior. My primary research focuses on the relationship between hippocampal aging and spatial cognition in homing pigeons. In addition to my work on old pigeons, I have studied the spatial-cognitive abilities of goldfish, whip spiders, and crayfish, as well as visual-attentional processes in humans.

Currently, I am working in collaboration with Drs. Verner Bingman (Bowling Green State University) and Anna Gagliardo (University of Pisa, Italy) to investigate possible Alzheimer's Disease-like pathology (specifically, congo-phillic amyloid plaques) in old homing pigeons.