My research interests are in three major areas: plant cell culture, anatomy and morphology of nonvascular and vascular plants, and evolutionary biology of plants.

My current research focuses on the production of useful plant metabolites from plant cell cultures derived from vascular cambial cells. These metabolites are diverse in chemical structure, vary with taxonomic groups of plants, and are used in applications ranging from medicine to cosmetology to agriculture. My work especially seeks to improve the cost efficiency and reliability of production of metabolites from plant cell cultures.

I also am interested in and have prior research projects in the use of anatomy, morphology, and nucleic acids to reconstruct patterns of evolution in plants, in particular, the mosses. This research helped to establish our current knowledge of broad relationships and trait evolution within mosses, an ancient lineage of plants. Related to this work, I also have research interests in the population biology of vascular plants, especially the pteridophytes (ferns and allies) and lycophytes (club mosses).