Leadership emergence theory discusses two pathways to leadership emergence—achievement (i.e. leaders’ behaviors) and ascription (i.e. leaders’ traits). Drawing from multilevel leadership emergence theory (Acton, Foti, Lord, & Gladfelter, 2019) which suggests that context influences the saliency of leadership emergence antecedents, our study simultaneously examined the incremental and relative importance of achievement and ascription antecedents to leadership emergence in contexts of low, medium, and high virtuality. In two independent samples—a laboratory experiment involving 86 teams (n = 340; sample one) and a semester long project involving 134 teams (n = 430; sample two)—we found that in low virtuality contexts, ascription factors accounted for incremental variance over achievement factors in predicting leadership emergence, and had larger relative importance. Conversely, in high virtuality contexts, achievement factors accounted for incremental variance over ascription factors in predicting leadership emergence, and had larger relative importance. Findings in medium virtuality contexts were mixed as achievement and ascription factors played relatively equal roles in the prediction of leadership emergence. Analyses employing other ratings of ascription (i.e. other-rated personality) found that a larger proportion of variance in leadership emergence was explained by other ratings than by self-ratings across all virtuality configurations.