Aim

Position of the alpine tree line ecotone around the globe corresponds to a growing season isotherm of approximately 6°C. Accordingly, tree lines are important indicators of Holocene global and regional environmental change. A central line of inquiry in tree line ecology is to better understand the mechanisms that dictate tree line position on the landscape, as well as the environmental conditions that cause upslope and downslope shifts in its position at centennial to millennial time‐scales. Here, we present a climate‐driven model to predict tree line position over the past 6,000 years.

Location

Sheep Mountain, located in the White Mountain Range of California, USA.

Time period

4750 bce to present.

Major taxa studied

Great Basin bristlecone pine (Pinus longaeva BK Bailey).

Methods

We use a climate‐driven tree line position model that utilizes a topoclimate raster surface of growing season average temperature to predict the spatial position and area of the alpine tree line ecotone across the mountain range. We then produce a time series of tree line position predictions at 500‐year intervals from 4750 bce to present, and compare the predictions to the growth dates and spatial locations of 61 remnant bristlecone pine samples from above modern tree line.

Results

The model indicates that tree line position in the White Mountains, CA migrated downslope throughout the Holocene until approximately 750 ce, rebounded slightly upslope by 1250 ce, and has since likely remained stationary. Applying the model under present‐day climatic conditions suggests the current tree line at Sheep Mountain may be out of climatic equilibrium by up to 250 vertical metres in some places.

Conclusion

The results support independent conclusions from global tree line analyses, underscore the temperature sensitivity of the tree line ecotone, and further develop our understanding of climate‐driven tree line dynamics.