Dr. Ryan is an associate professor in the Department of Biological Sciences at
Butler University. 

Articles

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Basking behavior of Emydid turtles (Chysemys picta marginata, Graptemys geographica, and Trachemys scripta elegans) in an urban landscape (with W. E. Peterman), Northeastern Naturalist (2009)

Basking is common in emydid turtles and is generally accepted to be in thermoregulatory behavior....

 

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Movement and habitat use of two aquatic turtles (with C A. Conner, B A. Douthitt, S C. Sterrett, and Carmen M. Salsbury), Urban Ecosystems (2008)

Our study focuses on the spatial ecology and seasonal habitat use of two aquatic turtles...

 

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Hydroperiod and metamorphosis in the small-mouthed salamander, Northeastern Naturalist (2007)

Ambystoma texanum (Small-mouthed Salamander) breeds primarily in temporary wetlands, and while natural history studies have...

 

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Legacy of land use in Southern Appalachian forests: effects on terrestrial salamander abundance along edges and within abandoned logging roads (with R. D. Semlitsch,; M. Chatfield; B. Drehman; K. Hamed; N. Pekarek; and M. Spath), Conservation Biology (2007)

Roads may be one of the most common disturbances in otherwise continuous forested habitat in...

 

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Descriptive ecology of a turtle assemblage in an urban landscape (with Conner C. A and B A. Douthitt), American Midland Naturalist (2005)

We studied turtle populations inhabiting a canal and a lake (both man-made) within a heavily...